Table 2.  Morphological data matrix. The character list is provided in Appendix 1.

 

Dendrobius          00000000000?0010000000000010000000000000000000000000000000000000000000000000000000000000000000000000000000000000000

Palophagoides       00000000000?0010000000000010000000000000000000000000000000000000000000000000000000000000000000000000000000000000000

Prionus             00000000000?0010000000000010000000000000000000000000000000000000000000000000000000000000000000000000000000000000000

Doydirhynchus       00010010000000110000000010000000?0000000010011100?0000000000000000000001100?000000000000000010120000000000000000001

Mecomacer           00010010000000110000000010000001?0000000010011100?0000000000000010000000100?000000000000000010120000000000000000?01

Rhynchitomacerinus  00010010000000110000000010000001?0000000010011100?0000000000000010000000100?000000000000000010120000000000000000?01

Ptychoderes         00010021000000?00000000?0000000010000000010011000000000000010000??01000110011000000000000101000100000000?0000000011

Toxonotus           00010021000000?00000000?0000000010000000010011000000000000010000??01000110011000000000000101000100000000?0000000011

Oxycraspedus        10010030000100000000000101100002100000000101122100010000000000000?000002000??100000000000000000100000000?0011000?01

Rhopalotria         10010030000100000000000101101002100000000101122100010000000000000?000002000??100000000000000000100000000?0011000?01

Rhinotia            10000000000100000000000101100002100000000001122100010000000000000?000000000??100000000000000100100000000?0011000?01

Apoderus            01000000000100000100000101?0?00210000100010102210001001000000000?1010001000010000?00?010000101110000000011101000011

Attelabus           01000000000100000100000101?0100210000100010102210001001000000000?1010001000010000?00?010000101110000000011101000011

Auletobius          01000000000100000100000101?0000210000100010112210001001000000000?1010001000000000000?010000101110000000011101000011

Merhynchites        01000000000100000100000101?0000210000100010112210001001000000000?1010001000000000000?010000101110000000011101000011

Caenominurus        001100100?01000001000?010110100000000100010112210011101010000000101110020001000100000000001000010010100001101000011

Antliarhinus        00010030100100000101?20101101102101?01000101122100111010100000001111101200011001000000111000100100101000?1101100111

Apion               0001003010010000010102010110110?101?010001?1122100111011100000001111101200011001000000111000100100101000?1101100111

Aporhina            0001002000010000010?010101101100110001000101122100111011100000001111101200010001000000111000100100101000?1101100111

Cylas               0001003000010000010?010101101100110001000101122100111010100000001111101200010001000000111000100100101000?1101100111

Ithycerus           0011002000010000010001010110110010000100011112?1011110100000000011111012000100010001?010100110010010100001101100111

Ocladius            001100300111000001000101011011121100010101111201?011101000000010111110?????110010?01?0101000100100101?00?1102100?11

Stromboscerini      001100300101?1000101031101101112110111011111122100111110000000001111101201111001110111101?010001001130001110211?011

Trigonotarsus       001100300101?1000101031101101112110111011111122100111110000000001111101201111001110111101?0100110011300011102112011

Rhinostomus         001100300101?1000101031101101112110111011111122100111110000000001111101201111001110111101?010011001130001110211?011

Rhynchophorus       001100300101?1000101041101101112110111011111122100111110000000001111101201111001110111101?0100110011300011102112011

Sitophilus          001100300111?1000101030101101112110111011111122100111110000000001111101201111001110111101?0100110011300011102112011

Sphenophorus        001100300101?1000101041101101112110111011111122100111110000000001111101201111001110111101?0100110011300011102112011

Stenopelmus         0011003001111000010103010110111211010101011112210011101000000000111110120211100100011010100000010010100011102110011

Lissorhoptrus       00110030011110000101030101101112110101011111122100111010000000001111101202111001000110101000?0010010100011102110011

Tanysphyrus         001100300111100001010301011011121101010101111221001110100000000011111012021110010?011010100000010010200011102110011

Penestes            0011003001111000010103010110111211010101011112210011101000000000111110120211100100011010100000010010100011102110011

Stenancylus         0011003001010000010103010110111211000101011112210011101000001000111111120111100211011010100010110120210111102111011

Cossonus            0011003001010000010103010110111211000101011112210011101000001000111111120111100211011010100000110120210111102111011

Araucarius          0011003001010000010103010110111211000101011112210011101000001000111110120111101210011010100010110120210111102110011

Phylloplatypus      0011003001010000010103010110111211000101011112210011101000001000111110120111101210011010100010110120210111102110011

Dendroctonus        001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100010110120210111102111011

Liparthrum          001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100010110120210111102111011

Phloeotribus        001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100010110120210111102111011

Hylurgonotus        001100300101000001010301011011121100011101110111001110100110?000111110120111101110011010100010110120210111102111011

Hylastes            0011003001010000010103010110111211000111011101110011101001101000111111120111101110011010100010110120210111102111011

Hylurgops           001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100010110120210111102111011

Ips                 001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100000010120210111102111011

Pityophthorus       001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100010110120210111102111011

Pityogenes          001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100000?10120210111102111011

Trypodendron        001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100000110120210111102111011

Dryocoetes          001100300101000001010301011011121100011101110111001110100110?000111111120111101110011010100000?10120210111102111011

Scolytus            0011003001010000010103010110111211000111011101110011101001100000111111120111100111011010100010110120210111102111011

Platypus            0011003001010100110103110110111211001111011101110011101000100000111112120111100110111000100000010?20211111102110011

Austroplatypus      00110030010101001101030101101112110011110111011100111010001000001111121201111001101110001000?0010?20211111102110011

Notoplatypus        0011003001010?00110103?101101112110011110111011100011010001000001111121201111001101110101000?0010?20211111102110011

Chaetastus          0011003001010?001101031101101112110011110111011100011010001000001111121201111001101110101000?0010?20211111102110011

Ampeloglypter       0011003001011000010103010110111211000101011112210011101000000101111110120111100101011010100010110120210111102110011

Sibariops           0011003001011000010103010110111211000101011112210011101000000101111111120111100101011010100010110120210111102110011

Odontocorynus       0011003001011000010103010110111211000101011112210011101000000101111111120111100101011010100010110120210111102110011

Callirhopalus       0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Diaprepes           0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Cylydrorhinus       0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Leptopius           0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Eudiagogus          0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Exophthalmus        0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Naupactus           0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Otiorhynchus        0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Asynonychus         0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Lachnopus           0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Polydrusus          0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Sitona              0011113001111000010103010111111211000101011112011011101000000000111110120111100100011010100010111120210111102110011

Tanymecus           0011113001111000010103010111111211000101011112011111101000000000111110120111100100011010100010111120210111102110011

Chrysolopus         0011003001111000010103010110111211010101011112210011101000000000111110120111100100011010100010111120210111102110011

Talaurinus          0011003001111000010103010110111211000101011112210011101000000000111110120111100100011010100010111120210111102110011

Gonipterus          00110030?11110000101130101101112110?0101011112210011101000000000111110120111100100011010100010111120210111102110011

Bronchus            0011?030?1?110000101?301011?1112110?0101011112210?11101000000000111110120111100100011010100010111120210111102110011

Listronotus         00110030?11110000101130101101112110?0101011112210011101000000000111110120111100100011010100010111120210111102110011

Oxyops              00110030?11110000101130101101112110?0101011112210011101000000000111110120111100100011010100010111120210111102110011

Listroderes         00110030011110000101130101121112110?0101011112210011101000000000111111120111100100011010100010111120210111102110011

Hypera              00110030?1011000010103010110111211000101011112210011101000000000111110120111100100011010100010110120210111102110011

Anthonomus          0011003011011000010103010110111211000101011112210011101000000000111110120111100102011010100010110120210111102110011

Curculio            0011003011011000010103010110111211010101011112210011101000000000111110120111100102011010100010010120210111102110011

Camarotus           00110030110110000101030101101112110?01000111122100111010000000?0111110120111100100011010100010110120210111102110011

Tachygonus          00110030110110000101030101101112110?0100011112210011101000000010111110120111100100011010100010110120210111102110011

Auletes             0011003011011000010103010110111211000101011112210011101000000111111110120111100100011010100010110120210111102110011

Rhinoncus           0011003011011000010103010110111211000101011112210011101000000111111110120111100100011010100010110120210111102110011

Phytobius           0011003011011000010103010110111211000101011112210011101000000111111110120111100100011010100010110120210111102110011

Araucarietius       00110030010110000101030101101112110?0101011112210011101000000000111110120111100100011010100010110120210111102110011

Eisingius           00110030010110000101030101101112110?0101011112210011101000000000111110120111100100011010100010110120210111102110011

Perelleschus        00110030010110000101030101101112110?0101011112210011101000000000111111120111100100011010100010110120210111102110011

Gymnetron           0011003011011000010103010110111210000101011112210011101000000000111110120111100100011010100010110120210111102110011

Myrmex              00110030010110000101030101101112110?0101011112210011101000000000111110120111100102011010100010110120210111102110011

Rhyephenes          0011003001011000011103010110111211000101011112210011101000000020111111120111100101011010100010110120210111102110011

Pseudomopsis        0011003001011000011103010110111211000101011112210011101000000020111111120111100101011010100010110120210111102110011

Bagous myriophyllae 0011003001111000010103010110111211010101011112210011101000000010111110120111100101011010100000110120210111102110011

Bagous americanus   0011003001111000010103010110111211010101011112210011101000000010111110120111100101011010100000110120210111102110011

Bagoini             0011003001111000010103010110111211010101011112210011101000000010111110120111100101011010100000110120210111102110011

Lixus               0011003001011000010103010110111211000101011112210?11101000000000111110120111100101011010100010110120210111102110011

Cholus              0011003001011000010103010110111211010101011112210011101000000000111111120111100101011010100010110120210111102110011

Conotrachelus       0011003001011000010103010110111211010101011112210011101000000010111111120111100101011010100010110120210111102110011

Epistrophus         0011003001011000010103010110111211010101011112210011101000000000111111120111100101011010100010110120210111102110011

Marshallius         0011003001011000010103010110111211010101011112210011101000000000111111120111100101011010100010110120210111102110011

Calvertius          0011003001011000010103010110111211010101011112210011101000000000111111120111100101011010100010110120210111102110011

Tranes              0011003001011000010103010110111211010101011112210011101000000000111110120111100101011010100010110120210111102110011

Pachylobius         0011003001011000010103010110111211010101011112210011101000000000111111120111100101011010100010110120210111102110011

Piazurus            00110030010110000101030101101012110?0101011112210011101000000111111110120111100111011010100010110120210111102110011

Macrocopturus       00110030010110000101030101101012110?0101011112210011101000000111111110120111100111011010100010110120210111102110011