#NEXUS 
[MacClade 3.01 registered to Jonathan Geisler, American Museum of Natural History]

BEGIN DATA;
DIMENSIONS  NTAX=39 NCHAR=123;
FORMAT MISSING=? GAP=-  SYMBOLS= " 0 1 2 3";
OPTIONS  MSTAXA=POLYMORPH ;

CHARLABELS
	[1]	Subarcuate_Fossa	[2]	Inflation_Tegmen_Tympani	[3]	Anterior_Proc_Petr	[4]	Tensor_Tympani_Fossa	[5]	Promontorial_Groove
	[6]	Stapedial_Groove	[7]	Mastoid_exp	[8]	Stylomastoid_Foramen	[9]	Facial_Nerve_Sulcus	[10]	'Petrosal/Basioccpital'
	[11]	Ectotympanic_Bulla	[12]	Pachyo_Bulla	[13]	Lat_furrow_Bulla	[14]	Med_furrow_Bulla	[15]	Art_Bulla_to_Squamosal
	[16]	'Tymp/Exocc_contact'	[17]	Sigmoid_Process	[18]	Sigmoid_process_shape	[19]	EAM_Tympanic_Floor	[20]	External_Auditory_Meatus
	[21]	'Basioccipital_crests_(falcate_p'	[22]	Carotid_Foramen	[23]	'Post-Glenoid_Foramen'	[24]	PRESENCE_For_ram_sup.	[25]	POSITION_for_ram_sup
	[26]	Posit_For_Ovale	[27]	Alisphenoid_Canal	[28]	Ant_Open_Alar_Canal	[29]	Post_Open_Alar_Canal	[30]	Foramen_Rotundum
	[31]	Mastoid_Foramen	[32]	Posttemporal_canal	[33]	Preglenoid_process	[34]	Supraorbital_process	[35]	posorbital_bar
	[36]	'Contact_Front/Max'	[37]	Lacrimal_Tubercle	[38]	palatine_fissure	[39]	Post_margin_nares	[40]	Mandibular_Foramen
	[41]	condyle	[42]	Dentary_angle	[43]	'Upp_Inc_1-2'	[44]	Incisor_Alignment	[45]	Embrasure_pits
	[46]	P1_presence	[47]	P3_roots	[48]	P4_protocone	[49]	P4_paracone_height	[50]	P4_metacone
	[51]	M2_metacone	[52]	M1_parastyle	[53]	ling_cing_M2	[54]	Ectocingula	[55]	Hypocone
	[56]	Trigon_basin	[57]	'Paraconule_M2/'	[58]	'Metaconule_M2/'	[59]	Preparacrista	[60]	Postparacrista
	[61]	Postmetacrista	[62]	M3	[63]	toothcomb	[64]	lower_incisors	[65]	Canine_shape
	[66]	p1	[67]	p3_metaconid	[68]	p4_metaconid	[69]	lin_cing_molars	[70]	m1_parac
	[71]	'm1_parac/crist_posit'	[72]	m2_parac	[73]	'm2_parac/crist_pos'	[74]	m3_parac	[75]	'm3_parac/cris_posit'
	[76]	m1_metaconid	[77]	m2_metaconid	[78]	m3_metaconid	[79]	m1_m2_hypoconulid	[80]	protoconid
	[81]	protoloph	[82]	metaloph	[83]	ant_low_lophs	[84]	post_low_lophs	[85]	Reen_groove
	[86]	Talonid_basin	[87]	M3_Hypoconulid	[88]	dp4	[89]	Cerv_vert_length	[90]	'Art_Sacral/Illium'
	[91]	Number_Sacrals	[92]	Scapular_Spine	[93]	Entepicondyle	[94]	Entepi_foramen	[95]	Olecranon_process
	[96]	Olec._fossa	[97]	Proximal_radius	[98]	Distal_radius	[99]	Centrale	[100]	Hand_Symm
	[101]	Greater_troch	[102]	3rd_troch	[103]	Distal_Femur	[104]	Astragalar_Canal	[105]	Navicular_Facet
	[106]	Sustentacular_facet	[107]	Cuboid_facet	[108]	Pes	[109]	Proxi_astrag	[110]	'Nav/Cub_fusion'
	[111]	'm3/m4_fusion'	[112]	Digit_1	[113]	Digit_2	[114]	Digit_5	[115]	Unguventral
	[116]	Ungdorsal	[117]	Digestive_epithelium	[118]	Lumen	[119]	Lumen_diverticulation	[120]	Penile_Muscle
	[121]	Hair	[122]	sebac_glands	[123]	Primary_Bronchi
	;
STATELABELS  
	1  present absent,
	2  uninflated inflated hyperinflated,
	3  absent present,
	4  shallow 'circular_pit,_no_groove' 'circular_pit,_anterior_groove' long_groove_between_tegmen_tymp,
	5  present absent,
	6  present absent,
	7  exposed_externally not_exposed_posteriorly,
	8  complete incomplete,
	9  absent 'ant._wall_=_squamosal' ant_wall_mas_of_petr 'ant._wall_=_meat._tube_ect',
	10  present absent,
	11  ring 'medial_edge_=_bulla',
	12  absent present,
	13  absent present,
	14  absent median_notch_on_posterior_rim prominent_AP_oriented_furrow,
	15  'contact_broad,_with_medial_pgp' cirucular_facet_on_short_entogl contact_with_crest_of_entogleno contact_absent,
	16  absent present,
	17  absent present,
	18  thin_and_transverse_plate broad_and_flaring,
	19  absent present_but_short present_and_long,
	20  absent_or_shallow elongate,
	21  absent present,
	22  absent 'present,_paired' falcate_processes,
	23  'present,_squam,_post_to_pgp' 'present,_med_to_petsq_sut' absent,
	24  present absent,
	25  lateral anterolateral,
	26  'ant_to_gf,_pos_wall_as' 'med_to_gf,_pos_wall_as' 'medial_to_gf,_post_wall_pet' pos_to_gf,
	27  absent present,
	28  within_sphenoorbital_fissure posterior_to_sphenoorbital_fiss,
	29  'well-separated_from_foramen_ova' within_deep_groove_with_f.o.,
	30  absent present,
	31  present absent,
	32  present absent,
	33  absent present,
	34  absent present,
	35  absent 'present,_almost_complete' 'present,_complete',
	36  absent present,
	37  absent present,
	38  small large,
	39  anteriorover_canines between_P1_and_P2 posterior_to_P2,
	40  small large,
	41  below_level_of_dentition even_with_superior_aspect_of_de substantially_higher_than_denti,
	42  level_with_ventral_edge below_ventral_edge,
	43  present I1_and_I2_absent,
	44  transverse_arc 'longitudinal,_long_prema',
	45  absent present,
	46  absent 'present_one-rooted' 'present_two-rooted',
	47  three two,
	48  present absent,
	49  equal_to_paracone_M1 greater_than_twice_M1_paracone,
	50  absent present,
	51  subequal_to_paracone half_paracone highly_reduced,
	52  absent weak strong,
	53  present absent,
	54  present absent,
	55  absent present,
	56  broad somewhat_narrow very_narrow,
	57  absent present,
	58  absent present,
	59  mesiodistal mesiolabial mesiolingual,
	60  mesiodistal distolabial distolingual,
	61  mesiodistal distolabial distolingual,
	62  'present,_larger_than_M2' 'present,_approx._equal_M2' reduced absent,
	63  absent present,
	64  small_and_vertical tusklike,
	65  oval triangular,
	66  present absent,
	67  absent present,
	68  absent present,
	69  absent 'continuous,_mesial-distal',
	70  present absent,
	71  cusp_or_crest_lingual 'cusp_or_crest_ant.betw_pr-pa' 'cusp_or_crest_m-d,_buccal_si',
	72  present absent,
	73  cusp_or_crest_lingual 'cusp_or_crest_ant.betw_pr-pa' 'cusp_or_crest_m-d,_buccal_si',
	74  present absent,
	75  cusp_or_crest_lingual 'cusp_or_crest_ant.betw_pr-pa' 'cusp_or_crest_m-d,_buccal_si',
	76  present absent,
	77  present absent,
	78  present absent,
	79  absent present,
	80  subequal_to_talonid twice_height_of_talonid,
	81  absent present,
	82  absent present,
	83  absent present,
	84  absent present,
	85  proximal absent distal,
	86  'basin,_hy_and_ent_present' reduced absent,
	87  long reduced absent,
	88  resembles_m1 6_cusped,
	89  long short very_long,
	90  broad narrow absent,
	91  1 '2-3' 4 '5-6',
	92  'large_acromion,_overhangs_gleno' small_acromion 'large_acromion,_does_not_overha',
	93  wide narrow,
	94  present absent,
	95  short long,
	96  shallow deep,
	97  single_fossa two_fossae three_fossae,
	98  single_fossa split_into_scaphoid_and_lunate convex_to_flat_distinct_facets,
	99  present absent,
	100  mesaxonic paraxonic,
	101  inferior_to_head_of_femur equal_to_head_of_femur above_head_of_femur,
	102  present highly_reduced absent,
	103  wide narrow,
	104  absent present,
	105  convex 'saddle-shaped' trochleated,
	106  narrow wide,
	107  absent 'present-small' 'present-large',
	108  'mesaxonic-3' paraxonic 'mesaxonic-4',
	109  flat grooved deeply_grooved,
	110  unfused fused,
	111  absent present,
	112  unreduced reduced highly_reduced,
	113  unreduced reduced highly_reduced,
	114  unreduced reduced highly_reduced,
	115  curved_inferiorly straight,
	116  transversely_compressed 'broad,_symmetrical' 'broad,_asymmetrical',
	117  nonglandular composite discoglandular glandular,
	118  unilocular plurilocular,
	119  absent present,
	120  abundant not_so_abundant,
	121  abundant absent_or_sparse,
	122  present absent,
	123  two three,
;

MATRIX
Didelphis          00001100000?????0??00101?10??0100000010010000??00?01?1000011100?0?01000000000011000000??00100000000021000000000000003000000
Asioryctes         ???01100000?????0??00?0??10??0??10???00010010(12)000012110210??120??000000000000001000010000?????????0?????0??000000000???????
Leptictis          00000000200?????0??000000(01)1100010000100010000100010200121120120?00000000000000110000100000?1001010?02?1000?01?010?00???????
Arctocyon          00000000?0?????????00?00010??1001000100?100001000000001000000200?0000000000000000000100000?00000000?0001001?00000?01???????
Hyopsodus          0101000?(03)0?????????00?00?110?000000?1000?000010000000010110001000011000000000010000010000020000111?000?1010000?0?010???????
Phenacodus         ??00110?(03)01?????0?000?01?1110010000010002000010001000(01)10110111000011010101000010000010100?2?0001111020110(01)0010010111???????
Meniscotherium     0100000?(03)??????????00?0??1110110010?100?200001000?0(01)1110111111000001010101000000011110000020001111?02011000000010111???????
Heptodon           ??0111?1300?????0??00?01?1110??0010?1?002000010001020010110010000111010101000010111110000???110111102011101020020211???????
Hyracotherium      ?10111??(03)1??????0??10?01?1110110010010002000020001010010110001100011010101000010111110000031110111102011100020020211???????
Equus              010011000110?20010200011?3100????02?1?00200001?0010?1110??11111000110101010000101111100000311111221020101000200222111000000
Diacodexis         0??1000?3010?0??10(01)00000010??00100?10?0??00001000000000011000100000000000000000000001001001?1101211?(12)110212120020010???????
Archaeotherium     100101103?1000011?210?21?00??0110020000001000110100(01)0110112001000000000000000010000010010011111121?10010212120022210???????
Elomeryx           110111103010000110210011?00??0010110100021000210000101101111100000110000000000100000100100?111112111??1?2?1120010012???????
Hippopotamus       110?111031100031102100(01)1?30??0??0111000011000110000100100022210111000000000000100000(02)0010031111111112110212120020012111?11?
Hexaprotodon       11????1031100031??210(01)11?2???0??0001000011000100000000100010010110110000000000100000200100311111211112102121100?0012111????
Sus                000111100110020010210021?20??0110000100020000210010101101122210011010000000000100000100100211111211112102121200200120001001
Agriochoerus       000?0000111001011021000??10??001000?100010000(02)0001020(01)10001110??0101000000000000000010010??1010121??0210211120020010???????
Poebrotherium      00011100?01002011?100?0??00??001011100002000021000021110000001000001000000000000000000012011110121111210211120022210???????
Camelus            000111002110010111100001?00??00101110000201?0000010011100011110000010000000000000000000120311101221102102121201222112111001
Ovis               1001010?2110013010200001?10??00101101100201?0000000211100011111001110000000000000000100100211101211?12102121211222121111001
Tragulus           0?01000031100(12)011?100001?00??00000100100201?00100(01)0011100010(01)110010000000000000000001001002111012111(12)21021112112121(02)111?00?
Andrewsarchus      ???????????????????10?(12)???1?0???10????0???000100?0(01)0000100???0???0??????????????0000??0????????????????????????????????????
Eoconodon          ???1110?(03)1?????????1000??0101??0000?1?00000?0100000000111100120000000000000000000000?00?????00?01??????????????????????????
Hapalodectes       ??00000?(03)0?????????0010??110011010?1000??0001100?1?2111200000(01)??00?00010101000010000012??????0?0???????????????????????????
Dissacus           0101000????????????1??20101?1??01???????00??01000112(01)0020(01)00020000000010101000010000?12?????00?111????101?1?00????11???????
Ankalagon          ????????????????????????????????????????0?????00011210020000020000000010101100?10000212?????10?11?????0?1?1?00????11???????
Sinonyx            ???10?00101000001?210?2???1?11?010?01000?0000100001210020000120000000010101101010000212????????????????????????????????????
Pachyaena          ??????0?????????1?210?????????101???1?00000001(01)001(01)2100201000200000000101011110100002120001010111101(12)0(01)01011100(12)0011???????
Mesonyx            1101000?1110000?1?210020101011?010????0??0?001(01)000121102000003??0000001010111?0100002120011110111101101(01)101110020011???????
Synoplotherium     ??????0???1?00??1?210?(12)??0101100100?100??0000??0??1211020000020000?000101011??010000212?0?101011110110111011?0020011???????
Harpagolestes      ??????001?1000011?210?2??11?1??010??100????0(01)1100122110200000300?1??0010101?1??10000212?????10111????0?1101?1??????????????
Pakicetus          01?2(01)1102011011110110?11?00??1??00?????010???2?110200002000011???(01)0011010??11?0100000120???????????????????????????????????
Ambulocetus        ?11???????111(01)1?1?0?0?(12)???0??0???0?????1?0?11??110?0110200???1?????01??1111?110?0000012???????1?010?1010???11?0?0011???????
Remingtonocetus    ?112111?1111111111011?(12)1?00??0??00???00????11??1?????????????0????????????????0?0000?1??002?????????2?0????????????????????
Cross_Whale        1112111?21111211???11111?0???0???1???00110011(12)????????????????????001?????????0?????012?1??????????????????????????????????
Protocetus         ??1211102111121110011?11?00?????01????1?????11011011?102000001?????????1?1??????0000????110????????????????????????????????
Basilosaurus       12131111211???2????01?11?00???11011??0(12)110011211102??0020000?300000001?1?1?111??00000?2?12?21110??012??????2?(01)0220?????????
Balaenoptera       121311112111103011011121?00??01101?1??20?0????????????????????00????????????????0000????1?02110?01?????????????222?????1111
Tursiops           121311110111123011001121??0??01101?1??21????0?????????????????00????????????????0000????1?02110?01?????????????222??1111111
;
END;

BEGIN ASSUMPTIONS;
	OPTIONS  DEFTYPE=unord PolyTcount=MINSTEPS ;

	TYPESET * UNTITLED  = unord: 1 3-13 16-18 20-22 24-34 36-38 40 42-45 47-50 53-55 57-60 62-89 91-100 102-106 109-111 114-123, ord: 2 14-15 19 23 35 39 41 46 51-52 56 61 90 101 107-108 112-113;

END; 

BEGIN TREES; 

	TRANSLATE
		1	Didelphis,
		2	Asioryctes,
		3	Leptictis,
		4	Arctocyon,
		5	Hyopsodus,
		6	Phenacodus,
		7	Meniscotherium,
		8	Heptodon,
		9	Hyracotherium,
		10	Equus,
		11	Diacodexis,
		12	Archaeotherium,
		13	Elomeryx,
		14	Hippopotamus,
		15	Hexaprotodon,
		16	Sus,
		17	Agriochoerus,
		18	Poebrotherium,
		19	Camelus,
		20	Ovis,
		21	Tragulus,
		22	Andrewsarchus,
		23	Eoconodon,
		24	Hapalodectes,
		25	Dissacus,
		26	Ankalagon,
		27	Sinonyx,
		28	Pachyaena,
		29	Mesonyx,
		30	Synoplotherium,
		31	Harpagolestes,
		32	Pakicetus,
		33	Ambulocetus,
		34	Remingtonocetus,
		35	Cross_Whale,
		36	Protocetus,
		37	Basilosaurus,
		38	Balaenoptera,
		39	Tursiops
	;
	TREE outgroup_constraint =  [&R] (3,2,1,(4,7,5,6,8,9,11,24,25,27,28,29,31,33,34,35,36,37,38,39));
	TREE outgroup2constraint =  [&R] (1,(3,2,(4,7,5,6,8,9,11,24,25,27,28,29,31,33,34,35,36,37,38,39)));
	TREE latest_constraint =  [&R] (1,(3,2,(4,22,7,5,6,8,9,11,24,25,27,30,28,29,31,33,34,35,36,37,38,39)));
	TREE svp_parax =  [&R] (1,(2,(24,(3,(4,(((((9,8),6),7),5),((((((16,13),14),12),(((20,17),18),21)),11),(23,(22,((((((31,29),30),28),(26,25)),27),(33,(34,(35,(36,(37,(39,38))))))))))))))));
	TREE adams =  [&R] (1,2,3,(24,(5,(6,(7,((8,9),10)))),(((17,(11,(12,(16,13,((14,15),((18,(19,(20,21))),(32,(33,(34,(35,36,(37,(39,38)))))))))))),23,22),4,(25,27,(26,(28,(30,(29,31))))))));
	TREE  * UNTITLED =  [&R] (1,2,(4,(3,((5,(6,(7,(8,9,10)))),((11,(17,(((12,16,13),(14,15)),(18,(19,(20,21)))))),(23,(22,((24,(25,(26,(27,(28,(30,(29,31))))))),(32,(33,(34,(35,(36,(37,(38,39)))))))))))))));

END;



BEGIN MACCLADE;
v 3.0
1661722971
0100&/0
0
0
END;