To acknowlege this copyright statement, please click the link below it.

Annual Reviews Copyright statement: "I am pleased to provide you complimentary one time access to my Annual Reviews article as a PDF file for your own personal use. Any further distribution or multiple distribution, publication or commercial usage of this copyrighted material would require submission of a permission request addressed to the Annual Reviews Permissions Department, email permissions@AnnualReviews.org."

I have read this copyright statement.