Evolutionary Biology Spring 2017
Meeting Time: Tuesdays and Thursdays, 9:30-10:45 am in TLS 154
Textbook: Futuyma, D.J. 2013. Evolution. 3rd ed. Sinauer Associates, Inc. (ISBN 978-1-60535-115-5)
Assigned readings are indicated below. Please read assigned chapters prior to coming to class.
Goal:The goal of this class is to teach the basic principles of evolutionary biology and the history of life on earth.
This website contains information for the lecture portion of the course only.
Please see HuskyCT for materials for the W portion of the course.
Check here for course announcements.
Dr. Janine N. Caira (Lectures 1–12)
Office: TLS 483
Office hours: by arrangement
Dr. Chris Simon (Lectures 13–25)
Office: Biopharm 305D
Office hours: by arrangement
Note: All emails must contain "EEB2245" in the subject line to avoid being filtered out and deleted
|Veronica Bueno||Katherine Nazario|
|Office: TLS 478||Office: BioPharm 323A|
|Phone: 486-1882||Phone: 486-3947|
|Students with last names A–K||Students with last name L–Z|
EEB 2245: Each half of the course counts for 50% of your lecture grade. Your grade in the first half of the course will be based on your performance on two lecture exams (for a total of 200 points). Your grade in the second half of the course will be based on your performance on three lecture exams (180 points total) and weekly, short on-line quizzes (20 points) (for a total of 200 points). However, your grade for Exams1–4 will be dropped. Please note: because you are allowed to drop one of these grades, we will not give make-up exams.
EEB 2245W: For W students your final grade in the lecture portion of the course will be calculated as above. This grade will constitute 75% of your final course grade. The W portion of the course, as determined by your “W” instructor, will constitute the remaining 25% of your final course grade. Please note that an F in either the W or lecture portion of the course will result in an F for the entire course. Dr. Paul Lewis (TLS 162; tel: 486–2069; email@example.com) is the coordinator of the W portion of the course. Refer to HuskyCT for further information on the W portion of the course. You will be assigned your W instructor at the mandatory library session you attend for the W portion of the course. You must sign up for a library session in HuskyCT by January 19th.
Please read assigned chapters, as indicated below, prior to class
This schedule is subject to change. Check regularly for updates!
|PART I: EVOLUTIONARY PATTERNS (CAIRA)|
|1||T Jan 17||Introduction to the Geological Time Scale; the Fossil Record||Ch. 4 (pp. 77–81)|
|2||Th Jan 19||Life in the Precambrian; evolution of the Metazoa||Ch. 5 (pp. 103–110)|
|3||T Jan 24||Cambrian explosion & Life in the Paleozoic||Ch. 5 (pp. 111–119)|
|4||Th Jan 26||Life in the Mesozoic||Ch. 5 (pp. 119–125)|
|5||T Jan 31||Life in the Cenozoic||Ch. 5 (pp. 125–132)|
|6||Th Feb 2||Evolution of primates||Ch. 4 (pp. 90–95)|
|T Feb 7||EXAM #1 (COVERS LECTURES 1–6)|
|7||Th Feb 9||Evolution of biodiversity & extinction||Ch. 7|
|8||T Feb 14||Characters, homology & homoplasy||Ch. 3 (pp. 51–63)|
|9||Th Feb 16||Systematics & reconstructing evolutionary history||Ch. 2|
|10||T Feb 21||Evolution and development||Ch. 3 (pp. 63–66) & Ch. 21|
|11||Th Feb 23||Biogeography & major patterns of distribution||Ch. 5 (pp. 129–132) & Ch. 6|
|12||T Feb 28||Continental drift & Historical Biogeography||Ch. 6|
|Th Mar 2||EXAM #2 (COVERS LECTURES 7–12)|
| Part II: EVOLUTIONARY PROCESSES (SIMON) |
See HuskyCT for weekly quizzes, periodic updates that track progress & introduce late breaking topics.
|13||T Mar 7||The uses of evolutionary biology. The evidence for evolution. The biblical literalists and the evolution debate; Refuting Creationist arguments. The Monkey Trial||Ch. 1 (pp. 1–3) and Ch. 23 (pp. 646–655)|
|14||Th Mar 9||Importance of variation. Sources of phenotypic variation. Variation due to the environment,phenotypic plasticity, common gardens. The Hardy-Weinberg equation: Why do we care? Is variability rare in natural populations?||Ch. 9 (to p. 235); Review Chapter 8 (basic genetics) on your own, especially pp. 208–209, mutation as a random process.|
|T Mar 14||SPRING BREAK|
|Th Mar 16||SPRING BREAK|
|15||T Mar 21||Epigenetic inheritance, variation in natural populations, Lewontin & Hubby, understanding deviations from HW ratios, effects of non-random mating, Inbreeding. Conservation biology and purging.||Ch. 9 remainder.|
|16||Th Mar 23||The Erosion of Genetic Variability by inbreeding. Genetic drift, consequences for conservation biology, Effective population size. Bottlenecks, founder events, gene flow models; gene flow studies. Geneflow x selection x drift.||CH. 10|
|17||T Mar 28||The interaction of ecology, climate, gene flow, bottlenecks. Northern purity/southern richness. Neutral theory.||Chs. 10 & 11|
|18||Th Mar 30||Introduction to selection, directional selection and the environment. Competitive character displacement, runaway sexual selection, the importance of heritability, multiple niche polymorphisms, the speed of directional selection, deleterious dominants.||Chs. 11 & 12|
|19||T Apr 4||Mutation and selection; fluctuating, frequency dependent, disruptive selection, Wright’s shifting balance, balancing selection. Cystic fibrosis & typhoid fever, Levels of selection. Coevolution, the evolution of virulence. Interdemic selection, Group selection, Kin selection, Cooperation, Inclusive fitness.||Chs. 11 & 12|
|Th Apr 6||EXAM #3 (COVERS LECTURES 13–19)|
|20||T Apr 11||Introduction to geographic variation. Character variation, parallel and perpendicular clines, geographic races of rat snakes and cicadas. Causes for clines. Human variation and the concept of Race. Ring species. Factors that inhibit gene flow- pre-mating isolation: Mating colors & dances, aggressive mimicry. Mating songs.||Chs. 9 (again), 17 & 18.|
|21||Th Apr 13||Post-mating and pre-zygotic: lock & key vs sexual selection and cryptic female choice, sperm-egg contact, natural selection at the molecular level. Post-mating and post-zygotic reproductive isolation, sterile hybrids. Post-mating and post-zygotic reproductive isolation (cont'd.), parasitic sterility, polyploids.||Ch. 17|
|22||T Apr 18||Speciation: The origin of biodiversity. Species concepts: typological, biological, phylogenetic, cohesion. Practical considerations: large biodiversity studies versus detailed studies of specific genera.||Chs. 17 & 18|
|23||Th Apr 20||Speciation Mechanisms: categories of speciation mechanisms. Butlin’s 2008 Table of speciation events over time. Allopatric, peripheral isolates, founder event speciation, reproductive character displacement, reinforcement. Periodical cicada case study. Allochronic speciation, contact zones, reproductive character displacement.||Chs. 17, 18 & 19 pp. 531–533.|
|24||T Apr 25||Speciation Mechanisms (cont.): Allochronic speciation, contact zones, reproductive character displacement (cont'd). Parapatric speciation, speciation with gene flow, North American chipmunk examples, sympatric speciation; host races. Translocations, polyploidy, polyphyletic species of Hyla.|
|25||Th apr 27||Parallel speciation. Chromosomal speciation with hybridization; co-adapted allele complexes. Sunflowers.||Read: Chs. 17, 18 & 19 pp. 531–533.|
|Final Exam Week EXAM #4 (COVERS LECTURES 20–25) & EXAM #5* (COVERS LECTURES 13–25)|