Evolutionary Biology Spring 2015
Meeting Time: Tuesdays and Thursdays, 9:30-10:45 am in TLS154
Textbook: Futuyma, D.J. 2013. Evolution. 3rd ed. Sinauer Associates, Inc. (ISBN 978-1-60535-115-5)
Objectives: The objectives of this course are to familiarize students with the mechanisms of evolutionary change (processes of evolution), major patterns of evolution, and the history of the diversity of life.
This website contains information for the lecture portion of the course only.
Please see HuskyCT for materials for the W portion of the course.
Problem Set 2 for Exam 2 is now available
Answer key for problem set 2.
This study guide provides some additional information about what to expect on exam 2. Use it to supplement your other course resources.
Exam 2 Review Session Tuesday March 3rd 7-9 PM BPB 130
Activity 12: Sharing game: what's good for the individual vs. what's good for the group. If you weren't able to play the sharing game, have a look to see what was learned!
Activity 11: Testing hypotheses of sexual selection. We began this in lecture. It is due at the beginning of class Thursday, Feb. 26. The additional data needed to complete the activity are here
Group game: the evolution of sharing. Come play! (This will count as activity 12).
All games will meet in TLS 313. There are 6 opportunities to play.
Activity 10: effects of dominant vs. recessive alleles. This is due online Tuesday, Feb. 24 by 9:15 am.
Activity 9 is the mean fitness table from Activity 8. Complete this table using the values from your Activity 8 simulations, and submit it at the beginning of class on Thursday, Feb. 19. Questions about Activity 9? Hopefully this will answer them!
Activity 8 is now available. This online activity simulating the effects of natural selection is due by Tuesday, February 17 at 9:15 am.
Problem Set 1 for Exam 1 is now available.
Answer key for problem set 1. Large file size: downloading may be slow.
Honors Conversion Students: More details on the different honors conversion options are now available here
Students interested in adding EEB2245W (or switching from EEB2245 to EEB2245W) should fill out this form and attend lecture. Those who are selected will be provided a permission number and asked to enroll within 24 hours. Unfortunately, it will not be possible to accommodate all students interested in the W version of this course.
SNOW DAY JAN 27TH: Make sure you check the updated syllabus and the new date for Exam 1
Part I, 20 January - 5 March
Dr. Elizabeth Jockusch
Office: PBB 305B
Office hours: 11-12 Tuesdays or by appointment
Part II, 10 March - 7 May
Dr. Chris Simon
Office: PBB 305D
Office hours: Anytime by appointment
Note: All emails must contain "EEB2245" in the subject line to avoid being filtered out and deleted
|James Bernot||Veronica Bueno||Geert Goemans|
|Office: TLS 478||Office: TLS 478||Office: PBB 323|
|Phone: 486-1882||Phone: 486-1882||Phone: 486-3947|
|Last names A-F||Last names G-N||Last names O-Z|
|Office hours: by appointment||Office hours: by appointment||Office hours: by appointment|
|Exam 1||50 points|
|Exam 2||100 points|
|Exam 3||50 points|
|Exam 4||100 points|
|Comprehensive Final Exam||70 points|
|Total Lecture Points||400 points|
EEB 2245W Grading: Your grade in the lecture portion of the course will be calculated as above. This grade will constitute 75% of your final course grade. Your grade in the W part of the course, as determined by your W instructor, will constitute the remaining 25% of your final course grade, except that an F in the W part of the course will result in an F for the entire course. An F in the lecture part of the course will also result in an F for the entire course.
- Arrive on time and stay until the end. If you must come late or leave early, sit by a door and leave as quietly as possible.
- Turn cell phones OFF and store them out of sight.
- Use laptops only for taking notes or other activity directly related to class.
- Recording is prohibited without the written permission of instructors.
- Course materials are the intellectual property of the course instructors. Students may not make these materials (including handouts, exams and activities) available electronically.
Activities: During the first half of the semester, there will be opportunities to earn points from a mixture of in-class and out-of-class activities. Each will be worth 3 points. A minimum of 13 opportunities will be available and the best 10 will count towards the final grade. Out-of-class activities must be submitted by the specified deadline for credit. No late assignments will be accepted. In-class activities must be submitted during the class period in which they take place. No make-ups will be given.
Missed exams: Any student who misses an exam without advance permission will receive a 0 for the exam. Permission to miss an exam requires, but is not guaranteed by, verifiable written documentation of the reason. A student who receives permission to miss an exam will have his or her grade for the missed work prorated based on his or her performance on the remainder of the exams. We will not give make-ups. Every student must take the final exam (and exam 4) during the scheduled final exam period unless permission to reschedule is obtained through the Dean of Students Office
Academic integrity: Plagiarism and cheating are violations of the student conduct code, and may be punished by failure in the course or, in severe cases, dismissal from the University. For more information, see Appendix A of the Student Conduct Code
Disabilities: If you have a disability for which you may be requesting an accommodation, you should contact a course instructor and the [www.csd.uconn.edu Center for Students with Disabilities] (Wilbur Cross Building, Room 201) within the first two weeks of the semester.
Please read assigned chapters, as indicated below, prior to class
This schedule is subject to change. Check regularly for updates!
|Date||Topic||Readings||Study Questions / Problem Sets|
|Part I: Jan 20 - Mar 6, Dr. Elizabeth Jockusch|
|Jan 20||Class organization; Introduction to the study of evolutionary biology||Ch 1||Study Questions
Activity 1, Evolutionary vs. non-evolutionary change
|Jan 22||Variation and the Hardy-Weinberg equilibrium||pp 217-228|| Study Questions|
Activity 2, testing for HWE
|Jan 27||SNOW DAY - CLASS CANCELED|
|Jan 29||Mutation and non-random mating||pp 189-208, 229-35, 247-8|| Study Questions |
More about kuru and prions
|Feb 3||Sampling effects and migration||Ch 10|| Study Questions |
Problem Set 1
|Feb 5||Conservation applications of population genetics||--||Study Questions|
|Feb 10||EXAM 1 (50 pts) & Natural selection||Ch 11|
|Feb 12||Evidence for natural selection||Ch 11||Study Questions|
|Feb 17||Genetics of natural selection||Ch 12||Study Questions|
|Feb 19||Modes of natural selection||Ch 12 & 13||Study Questions|
Activity 9-Mean fitness table at end of Activity 8 due in class today
|Feb 24||Sexual selection and female choice||Ch 15||Study Questions
Activity 10-Selection on dominant vs. recessive alleles : online portion due before class; bring your graphs to class
|Feb 26||Genetic conflict and levels of selection||Ch 16||Study Questions|
|Mar 3||Geographic variation and speciation||pp 483-491||Study Questions
Review Session tonight 7-9pm BPP 130
|Mar 5||EXAM 2 (100 pts, midterm covering all lectures)|
|Part II: Mar 10 - May 7, Dr. Chris Simon|
|Mar 10||Mechanisms of Speciation (taught by Dr. Jockusch)||Ch 17 & 18||Study Questions|
|Mar 12||Species Concepts. Hybridization (taught by Dr. Wade)||Ch 17 & 18||Discussion Questions|
|Mar 17||SPRING BREAK|
|Mar 19||SPRING BREAK|
|Mar 24||Speciation Mechanisms. Spatial, temporal, chromosomal with and without hybridization, parasitic.|| Discussion Questions |
|Mar 26|| Systematics, the study of biodiversity and its origins.
Problems in constructing relationships: polymorphisms and homoplasy.
|Ch 2 & 3|| Discussion Questions|
Lecture 16 part I
Lecture 16 part II
|Mar 26||Discussion Session 7-8pm BPB 131|| Discussion questions
Lectures 14, 15 & 16
|Mar 30||Discussion Session 8-9pm BPB 131|| Discussion questions
Lectures 14, 15 & 16
|Mar 31||Homoplasy (continued): convergence, parallelisms, and reversals in evolution.||Ch 2 & 3||Discussion Questions|
|Apr 2|| Reconstructing evolutionary trees from morphological and molecular data.
How molecules evolve. Is there a molecular clock?
|Ch 2 & 3|| discussion questions|
|Apr 2||Discussion Session 7-9pm BPB 131|| Discussion questions
Lectures 17 & 18 + Exam review
|Apr 7||Discussion Session 7-9pm BPB 131|| Discussion questions
Lectures 17, 18 & 19 + Exam review
|Apr 7|| Overview of life continued. The origin of evolutionary novelties: Body plans, constraints; pre-adaptation, modification of existing traits:
gene duplication, gene regulation
|Ch 21 & 22||discussion questions |
|Apr 9|| A review of the tree of life and the major innovations in animal evolution.
EXAM 3 (50 pts)
|Apr 9||Discussion Session 7-8pm BPB 131|| Discussion questions
Lectures 19 & 20
|Apr 13||Discussion Session 8-9pm BPB 131|| Discussion questions
Lectures 19 & 20
|Apr 14||Evolutionary novelties (continued) Homeobox genes, Master control genes. Flies with eyes on their wings. Ontogeny and phylogeny, Allometry.||Ch 21 & 22||discussion questions|
|Apr 16|| Introduction and overview of the fossil record. The origin of life.
The RNA world. Prokaryote world. The origin of animals; the Ediacaran Fauna. Mass extinctions.
|Ch 4 & 5, pp 168-171, Box 7A||discussion questions|
|Apr 16||Discussion Session 7-8pm BPB 131|| Discussion questions
Lectures 21 & 22
|Apr 20||Discussion Session 8-9pm BPB 130|| Discussion questions
Lectures 21 & 22
|Apr 21||The Paleozoic: Cambrian explosion (or was it?). The origin of vertebrates and the invasion of land. Ordovician (the age of jawless vertebrates), Silurian (first life on land), Devonian (the age of fishes). Carboniferous (Dragonflies w/ 2 ft. wing span, clubmoss forests); Permian. The origin of mammals. The Permo-Triassic boundary mass extinction.||Ch 4 & 5, 168-171, box 7A||discussion questions|
|Apr 23||The Mesozoic: The age of reptiles. Pangea breaks up followed by Laurasia and Gondwanaland. The evolution of birds from dinosaurs, insects and angiosperms radiate. The K-T Boundary. The extinction of the dinosaurs. Birds and mammals cross the boundary.||Ch 4, 5 & 6||discussion questions|
|Apr 23||Discussion Session 7-8pm BPB 131|| Discussion questions
Lectures 23 & 24
|Apr 28||The Cenozoic: Greenhouse to Icehouse. Continental drift, land bridges, mountain building. Modern biogeographic distributions take shape. The great American interchange. Primate evolution.||Ch 4, 5 & 6||discussion questions|
|Apr 30||Human evolution; Mitochondrial Eve and her relatives. Africa, our most diverse continent. Humans invade Asia and the Pacific and later North America. Biogeography and Biodiversity.||Ch 4, 5 & 6||discussion questions|
|May 7||EXAM 4 (100 pts) & COMPREHENSIVE FINAL EXAM (70 pts)|