Evo Devo Fall2013

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EEB 5333, Fall 2013
Evolutionary Developmental Biology
Meeting Time: Tuesdays and Thursdays, 2:00-3:15 pm, Bamford Room (TLS 171B)

Contents

Description

This is an advanced course that explores the interface between evolutionary biology and developmental biology. In addition to considering how developmental pathways evolve, and the developmental basis of phenotypic evolution, we will ask what novel insights emerge from a synthesis of these fields. Major topics to be considered include the following: developmental constraints, homology, plasticity, novelty and evolvability.

Instructors

Dr. Elizabeth Jockusch
Office: Biology/Pharmacy 305B
Phone: (860) 486-4452
Office hours: by appointment

Dr. Carl Schlichting
Office: Torrey Life Sciences 366
Phone: (860) 486-4056
Office hours: by appointment


Announcements and Handouts

Books & Sources in Evolutionary Developmental Biology


Expectations and Grading

This course has a mixed lecture/discussion format. In general, Tuesdays will be used for lectures that provide an overview and background information. Thursdays will be dedicated to student-led discussion based on reading from the literature. We expect everyone to participate actively in the class. In order to help prepare for discussions, you should write a brief (<1 page) reaction piece to each set of readings, highlighting your thoughts about the readings, connections between them or questions raised by them. This will be handed in each week. Responsibility for leading the discussions will rotate. 
 You are expected to complete an independent project on a topic of your choice. Ideally, your topic will be closely connected to your own research interests, and also integrate multiple topics covered in class . You should discuss your choice of topics with one of us no later than Friday, October 4. To ensure that you are on track, an outline and relevant references are due on Tuesday, Nov. 12. The final project paper is due Tuesday, Dec. 3, and the last week of class will be devoted to presentations and discussions based on these projects. You will be in charge of assigning a paper relevant to your presentation.
 There will be one take-home exam due the day of the Final Exam. It will consist of one or several essay questions that will ask you to integrate the knowledge that you have acquired during the course. 



Deadlines

WeeklyDiscussion pieces
Friday, Oct. 4Independent project topic approved
Tuesday, Nov. 12Independent project outline and references due
Tuesday, Dec. 3Independent project paper due
Dec. 3-5Independent project presentations
Tuesday, Dec. 10Final exam due


Course grade

Discussions35%, including participation, leading discussion, and reaction pieces
Individual project40%, based on presentation (15%) and paper (25%)
Final exam25%


Topics and Readings

Week 1 (Aug. 27/29): Overview of evolutionary developmental biology

Discussion Leader: Elizabeth
Discussion Readings:
Discussion Questions:

Week 2 (Sept. 3/5): Morphological and molecular patterns

Discussion Leader:
Discussion Readings: The developmental hourglass and the phylotypic stage
Discussion Questions:

Week 3 (Sept. 10/12): The molecular building blocks of development

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Week 4 (Sept. 17/19): Evolution of developmental networks

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Week 5 (Sept. 24/26): Development and homoplasy

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Week 6 (Oct. 1/3): Developmental constraints and evolution

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Week 7 (Oct. 8/10): Development and homology

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Week 8 (Oct. 15/17): Novelty

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Week 9 (Oct. 22/24): Modularity

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Week 10 (Oct. 29/31): Robustness and canalization

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Week 11 (Nov. 5/7): Plasticity

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Week 12 (Nov. 12/14): TBA

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Week 13 (Nov. 19/21): Evolvability

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Week 14 (Nov. 26/28): Happy Thanksgiving!

Week 15 (Dec. 3/5): Project Presentations

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