Andrew Bush

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Associate Professor of Paleobiology
Department of Ecology & Evolutionary Biology
and Center for Integrative Geosciences
University of Connecticut

Contact Information

Email: andrew.bush [at]
Office (EEB): Torrey Life Sciences 472
Lab: Torrey Life Sciences 469
Office (Geo): Beach 240
Phone (EEB): 860-486-9359
Phone (Geo): 860-486-9385
Fax: 860-486-6364

Research Interests (click for more info)

  • Changes in marine paleoecology and biodiversity
  • The Late Devonian (Frasnian/Famennian) mass extinction
  • Upper Devonian stratigraphy in the Appalachian Basin
  • Ecologic gradient analysis of fossil species
  • Hartford Basin paleontology


  • Paleobiology Seminar (various topics; EEB 5895/GSCI 6130)
  • Paleobiology (EEB/GSCI 4120)
  • Earth History and Global Change (GSCI 3010)
  • Principles of Biology II (BIOL 1108)


  • Bush, A. M., and R. K. Bambach. 2015. Sustained Mesozoic-Cenozoic diversification of marine Metazoa: a consistent signal from the fossil record. Geology, in press.
  • Bush, A. M., Csonka, J. D., DiRenzo, G. V., Over, D. J., Beard, J. A. 2015. Revised correlation of the Frasnian-Famennian boundary and Kellwasser events (Upper Devonian) in shallow marine paleoenvironments of New York State. Palaeogeography, Palaeoclimatology, Palaeoecology. 433:233-246.
  • Getty, P.R., Hardy, L., and Bush, A.M. 2015. Was the Eubrontes track maker gregarious? Testing the herding hypothesis at Powder Hill Dinosaur Park, Middlefield, Connecticut. Bulletin of the Peabody Museum of Natural History. 56:95–106.
  • Fichman, M.E., Crespi, J.M., Getty, P.R., and Bush, A. M. 2015. Retrodeformation of Carboniferous trace fossils from the Narragansett Basin, United States, using raindrop imprints and bedding-cleavage intersection lineation as strain markers. Palaios 30:574-588.
  • Bush, A. M., S. B. Pruss, and J. L. Payne, eds. 2013. Ecosystem Paleobiology and Geobiology. Paleontological Society Papers 19.
  • Bush, A. M., and S. B. Pruss. 2013. Theoretical ecospace for ecosystem paleobiology: energy, nutrients, biominerals, and macroevolution. In A. M. Bush, S. B. Pruss, and J. L. Payne, eds. Ecosystem Paleobiology and Geobiology. Paleontological Society Papers 19:1-20.
  • Bush, A. M., S. B. Pruss and J. L. Payne. 2013. Ecosystem paleobiology and geobiology: connecting the biological and earth systems. In A. M. Bush, S. B. Pruss, and J. L. Payne, eds. Ecosystem Paleobiology and Geobiology. Paleontological Society Papers 19:xi-xiii.
  • Getty, P. R., R. Sproule, D. L. Wagner, and A. M. Bush. 2013. Variation in wingless insect trace fossils: insights from neoichnology and the Pennsylvanian of Massachusetts. Palaios 28:243-258.
  • Christie, M., S. M. Holland, and A. M. Bush. 2013. Contrasting the ecological and taxonomic consequences of extinction. Paleobiology 39:538-559.
  • Getty, P. R., A. Judge, J. Csonka, and A. M. Bush. 2012. Were Early Jurassic dinosaurs gregarious? Reexamining the evidence from Dinosaur Footprint Reservation in Holyoke, Massachusetts. Pp. A1-A18 in M. A. Thomas, ed. Guidebook for Fieldtrips in Connecticut and Massachussets: Geological Society of America Northeast Section, 47th Annual Meeting, Hartford, Connecticut. State Geological and Natural History of Connecticut Guidebook 9.
  • Bush, A. M., and Novack-Gottshall, P. M. 2012. Modelling the ecological-functional diversification of marine Metazoa on geological time scales. Biology Letters, 8:151-155.
  • Bush, A. M., R. K. Bambach, and D. H. Erwin. 2011. Ecospace utilization during the Ediacaran Radiation and the Cambrian Eco-explosion. Pp. 111-133 in M. LaFlamme, J. D. Schiffbauer, and S. Q. Dornbos, eds. Quantifying the evolution of early life: numerical approaches to the study of fossils and ancient ecosystems. Springer, Dordrecht.
  • Bush, A. M., and R. K. Bambach. 2011. Paleoecologic megatrends in marine metazoa. Annual Review of Earth and Planetary Sciences 39:241-269.
  • Getty, P. R., and A. M. Bush. 2011. Sand pseudomorphs of dinosaur bones: Implications for (non-) preservation of tetrapod skeletal material in the Hartford Basin, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 302:407–414.
  • Bush, A. M., and R. I. Brame. 2010. Multiple paleoecological controls on the composition of marine fossil assemblages from the Frasnian (Late Devonian) of Virginia, with a comparison of ordination methods. Paleobiology 36:573–591.
  • Bush, A. M., and G. M. Daley. 2008. Comparative paleoecology of fossils and fossil assemblages. In P. H. Kelley and R. K. Bambach, eds. From evolution to geobiology: research questions driving paleontology at the start of a new century. Paleontological Society Paper 14:289-317.
  • Wang, S. C., and A. M. Bush. 2008. Adjusting global extinction rates to account for taxonomic susceptibility. Paleobiology 34:434–455.
  • Bush, A. M., R. K. Bambach, and G. M. Daley. 2007. Changes in theoretical ecospace utilization in marine fossil assemblages between the mid-Paleozoic and late Cenozoic. Paleobiology 33:76–97.
  • Bambach, R. K., A. M. Bush, and D. H. Erwin. 2007. Autecology and the filling of ecospace: key metazoan radiations. Palaeontology 50:1–22.
  • Bush, A. M., M. Kowalewski , A. Hoffmeister , R. K. Bambach, and G. M. Daley. 2007. Potential paleoecologic biases from size-filtering of fossils. Palaios 22:612–622.
  • Bush, A. M. 2006. Crime Scene Investigation—Permian. Review of Erwin, D. H. 2006. Extinction: How Life on Earth Nearly Ended 250 Million Years Ago. Science 311: 1868-1869.
  • Bush, A. M., and R. K. Bambach. 2004. Did alpha diversity increase during the Phanerozoic? Lifting the veils of taphonomic, latitudinal, and environmental biases in the study of paleocommunities. Journal of Geology 112:625–642.
  • Bush, A. M., M. J. Markey, and C. R Marshall. 2004. Removing bias from diversity curves: the effects of spatially organized biodiversity on sampling-standardization. Paleobiology 30:666–686.
  • Bush, A. M., M. G. Powell, W. S. Arnold, T. M. Bert, and G. M. Daley. 2002. Time-averaging, evolution, and morphologic variation. Paleobiology 28:9–25.