Frog and Toad Study Guide
Herpetology Home


Remember that you should be able to identify specimens to family by external morphological characteristics alone.  Do not rely upon coloration characteristics since these are often bleached out in preserved specimens.  Typical coloration patterns also tend to vary greatly in nature.  The following information was presented in lab at each station.

Anuran families

Connecticut anurans

Families of Frogs and Toads

Know specimens to family level and subfamily anurans.  Know Distribution and Representative Forms information contained in the Anura handout, even if there are no representative specimens.


Mesobatrachia
Neobatrachia
Neobatrachia
Bufonoidea
Ranoidea
       


Ascaphidae

- males have copulatory organ derived from the cloaca
- lack tympana
- outermost toe thicker than others

Specimens on display:
Ascaphus truei


Leiopelmatidae

No specimens available



Discoglossidae

- small to medium size
- disc-shaped tongue
- pupil vertically elliptical or triangular
- generalized semi-aquatic morphology

Specimens on display:
Alytes sp.
Discoglossus pictus



Bombinatoridae

- Bombina has toad-like morphology, Barbourula more aquatic
- distinctive patterns of black or green and bright red, yellow, or orange on the belly

Specimens on display:
Bombina bombina
Bombina orientalis
Bombina variegata




Pipidae

- aquatic morphology: webbing on back feet and flattened bodies
- pupil is round, eyes small and dorsal, eyelids poorly developed or absent
- hands with long fingers, only webbed in Hymenochirus and Pseudhymenochirus
- toes have distinct keratinous claw-like tips
- limbs splayed laterally
- some species have lateral-line system
- lack tongues

Specimens on display:
Pipa pipa
Xenopus laevis




Rhinophrynidae

- morphology adapted for burrowing, resembles North American microhylid frogs
- well-developed spade present on foot
- robust body
- head pointed with thickened, cornified skin at tip
- small eyes, vertical pupils
- lack tympanum
- tongue is free at the front, protruded from groove at front of mouth

Specimens on display:
Rhinophrynus dorsalis



Pelobatidae

- vertically elliptical pupils
- relatively short-legged
- large metatarsal tubercules on the hind feet for burrowing
- many have tuberculate skin with enlarged parotoid glands on dorsum
- large pectoral glands in Scaphiopus holbrooki

Specimens on display:
Pelobates fuscus
Scaphiopus couchii
Scaphiopus holbrookii
Spea bombifrons
Spea hammondii
Spea intermontanus




Megophryidae

- large terrestrial or fossorial frogs, similar to Pelobatidae
- very glandular skin, unusual clusters of large granular glands found in axillary, pectoral, and femoral region in both sexes
- some have fleshy dorsolateral folds
- Megophrys notable for cryptic coloration and disruptive body outlines
- Megophrys nasuta has projections of skin on snout and over eyes

Specimens on display:
Megophrys nasuta



Pelodytidae

No specimens available



Allophrynidae

No specimens available



Brachycephalidae

No specimens available



Bufonidae

- most have terrestrial morphology, a few have stream or arboreal morphology
- heavy and squat with short legs
- rough skin with conspicuous parotoid glands on head
- skull usually co-ossified with overlying skin
- no teeth
- pupil is horizontal
- Atelopus has thin body, long legs, and slender toes; lacks tympanum; some have aposematic coloration
- Melanophryniscus resembles Bufo, lacks cranial crests and parotoid glands

Specimens on display:

Atelopus sp.
Atelopus carrikeri
Atelopus muisca
Bufo
sp.
Bufo americanus
Bufo boreas
Bufo bufo
Bufo calamita
Bufo coniferus
Bufo fowleri (Bufo woodhousii fowleri)
Bufo granulosus
Bufo marinus
Bufo paracnemis
Bufo punctatus
Bufo quercicus
Bufo terrestris
Bufo valliceps
Bufo vertebralis
Bufo viridis

Melanophryniscus stelzneri



Centrolenidae

- arboreal morphology
- small and delicate
- pupil horizontal
- terminal phalanges T-shaped with expanded toe-discs
- nearly transparent skin on venter
- males have bony process on humerus

Specimens on display:
Centrolene prosoblepon



Heleophrynidae

No specimens available



Hylidae

- extremely variable in size and appearance, most have arboreal morphology
- most have claw-shaped terminal phalanges with toepads
- at least short webs between toes
- intercalary cartilage present
- many have co-ossification of skin on skull
- pupil horizontal except in phyllomedusines

Subfamily Hylinae

- most with arboreal morphology
- Pseudacris and Acris are less arboreal, little or no enlargement of the toes

Specimens on display:
Acris crepitans
Acris gryllus
Duellmanohyla rufioculis
Hyla andersoni
Hyla arenicolor
Hyla aurea
Hyla cinerea
Hyla ebraccata
Hyla femoralis
Hyla gratiosa
Hyla nana
Hyla pseudopuma
Hyla regilla
Hyla squirella
Hyla versicolor
Osteopilus septentrionalis
Phrynohyas venulosa
Pseudacris crucifer
Pseudacris nigrita
Pseudacris ocularis
Pseudacris ornata
Pseudacris triseriata
Pternohyla fodiens
Scinax boulengeri
Scinax fuscovaria
Smilisca baudini
Smilisca phaeota
Smilisca sordida


Subfamily Phyllomedusinae

- large treefrogs
- some have long legs with prehensile, monkey-like feet
- many have conspicuous glands on dorsum
- pupil vertically elliptical

Specimens on display:
Agalychnis callidryas
Agalychnis spurrelii
Pachymedusa dacnicolor
Phyllomedusa lemur


Subfamily Hemiphractinae

No specimens available

Subfamily Pelodryadinae

No specimens available



Leptodactylidae

- very diverse in morphology and size
- webs between toes often greatly reduced
- no intercalary cartilage
- pupil horizontal in most genera

Subfamily Leptodactylidae

- Leptodactylus has typical frog-like appearance, large range in size
- Rana-like species can be distinguished by having little or no webbing on hind feet
- in many species males have enlarged forearms and spines on the “thumbs” or chest

Specimens on display:
Leptodactylus albilabris
Leptodactylus bolivianus
Leptodactylus laticeps
Leptodactylus melanonotus
Leptodactylus mystacinus
Leptodactylus pentadactylus
Physalaemus biligonigerus
Physalaemus pustulosus


Subfamily Ceratophryinae

- squat, toad-like frogs with wide mouths
- some have skin flaps, bony projections, and camouflaging coloration
- large teeth in the upper jaw and bony projections from the lower jaw

Specimens on display:
Ceratophrys ornata

Subfamily Telatobiinae

No specimens available

Subfamily Eleutherodactylinae

- relatively slender with narrow heads, long legs, long toes, enlarged toe pads, and little or no webbing on feet
- some species like E. bombifrons are squat and toad-like with short legs and broad heads

Specimens on display:
Eleutherodactylus biporcaus
Eleutherodactylus bogotensis
Eleutherodactylus bransfordi
Eleutherodactylus bufoniformis
Eleutherodactylus caryophyllaceus
Eleutherodactylus coqui
Eleutherodactylus crassidigitus
Eleutherodactylus cruentus
Eleutherodactylus fleischmanni
Eleutherodactylus melanostictus
Eleutherodactylus ricordii
Eleutherodactylus vocator


Subfamily Cyclorhamphinae

No specimens available



Myobatrachidae

No specimens available



Pseudidae

- highly aquatic, extensive webbing on hind feet, hindlimbs robust
- elongate, ossified intercalary elements
- pupil horizontal, eyes large and protuberant
- Pseudis paradoxa has tadpole larger than adult

Specimens available:
Pseudis paradoxa



Rhinodermatidae

No specimens available



Sooglossidae

No specimens available



Dendrobatidae

- most are small in size with bright aposematic coloration
- pupil horizontal
- pair of dermal scutes on the dorsal surfaces of the fingers
- Colostethus cryptically colored

Specimens on display:
Colostethus inguinalis
Colostethus nubicola
Colostethus subpunctatus
Dendrobates
sp.
Dendrobates auratus



Arthroleptidae

- generally semi-aquatic morphology
- Trichobatrachus males have long hair-like skin projections during the breeding season

Specimens on display:
Arthroleptella hewitti
Trichobatrachus robustus



Hemisotidae

No specimens available



Hyperoliidae

- arboreal and terrestrial morphologies
- most are small
- pupil round, horizontal, or vertically elliptical
- some have toe-discs and claw-shaped terminal phalanges
- some have aposematic coloration

Specimens on display:
Hyperolius rossii



Mantellidae

No specimens available



Microhylidae

- diverse morphology, usually small in size
- body shape varies from squat and small-headed to globular and toad-like to arboreal morphology with expanded tips of digits
- prominent fold of skin across the head behind the eyes
- pupil of eye horizontal or round in most species

Subfamily Microhylinae

- burrowing morphology, triangular-shaped bodies with narrow heads

Specimens on display:
Gastrophryne carolinensis
Kaloula borealis
Uperodon
sp.

Subfamily Breviceptinae

No specimens available

Subfamily Asterophryninae

No specimens available

Subfamily Genophryninae

No specimens available

Subfamily Phrynomerinae

No specimens available

Subfamily Melanobatrachinae

No specimens available

Subfamily Cophylinae

No specimens available

Subfamily Dyscophinae

No specimens available



Ranidae

- size and body form extremely varied
- slender to robust, terrestrial to semi-aquatic morphology
- tongue bicornate at posterior
- pupil horizontal in most species
- Amolops and Staurois have enlarged toe pads for holding onto rocks in mountain streams, tadpoles have large sucker mouthparts
- Ceratobranchus is ornamented to resemble a dead leaf
- Palmatorappia has arboreal morphology

Subfamily Raninae

Specimens on display:
Afrana fuscigula
Rana aurora
Rana boylii
Rana cascadia
Rana catesbeiana
Rana clamitans
Rana esculenta
Rana grylio
Rana heckscheri
Rana nigromaculata
Rana palustris
Rana pipiens
Rana pretiosa
Rana ridibunda
Rana septentrionalis
Rana temporaria
Rana virgatipes
Rana warszewitschii
Strongylopus grayi


Subfamily Petropedetinae

No specimens available


Rhacophoridae

- variable in size, arboreal morphology with enlarged toe-discs
- pupil horizontal
- Rhacophorus wardtii has large webs on feet for gliding

Specimens available:
Unidentified species


Connecticut Frogs and Toads

Be able to identify anurans to genus and species level and be able to sex them.  Also know their taxonomy (family, subfamily). You do not need to know specific distributions within Connecticut.  Know natural history information given in the Anura handout.  

Pelobatidae
Bufonidae
Hylidae
Ranidae


Scaphiopus holbrookii (Eastern spadefoot toad)
Family Pelobatidae

Morphological characteristics:
- body broad and thick
- skin relatively smooth
- eyes large and protruding, vertical pupils
- distinct round parotoids
- pectoral glands
- no cranial crests
- distinct tympanum
- fingers with no webbing
- hindlimbs short
- two metatarsal tubercules (spades), outer one larger with cutting edge (sickle-shaped)
- feet with extensive fleshy webs

Living specimens:
- dorsal color brown, grayish, or greenish
- two yellow stripes forming a lyre- or hourglass-shaped pattern on back



Bufo americanus (American toad)
Family Bufonidae

Morphological characteristics:
- body broad and thick
- skin rough with many warts, 1-3 warts in each dark dorsal spot
- warts on shank often larger than those on thigh
- venter granular
- large protruding eyes with horizontal pupils
- large kidney-shaped (reniform) parotoids behind eyes
- distinct cranial crests, separated from parotoid or connected by spur
- hindlimbs short
- extensive webbing on hind feet

Living specimens:
- dorsal color brown, yellowish, gray, or brick red
- throat of male dark
- venter often mottled or spotted
- light middorsal stripe may be present



Bufo fowleri (Fowler’s toad)
Family Bufonidae

Morphological characteristics:
- similar body shape to B. americanus
- parotoids touch postorbital ridge of cranial crest, no spur connection
- parotoids oval
- usually 3-6 warts in each dark dorsal spot
- warts on shank not larger than those on thigh

Living specimens:
- color often greenish-gray with distinct dark spots
- venter usually unspotted
- usually light middorsal stripe



Pseudacris crucifer (Spring peeper)
Family Hylidae

Morphological characteristics:
- body narrow and thin
- horizontal pupils
- no parotoids
- hind limbs long
- small round discs on toes

Living specimens:
- tan, grayish, or brownish
- distinct X mark on back
- throat of males dark



Hyla versicolor (Gray treefrog)
Family Hylidae

Morphological characteristics:
- body stout
- skin slightly rough in appearance
- hind limbs long
- round disks on toes

Living specimens:
- color gray or light green (color can change depending on light and temperature)
- pattern resembling lichens on tree bark
- bright orange wash on rear of thighs and groin
- black-bordered light area below eye
- rear of thigh often with a dark bar
- throat of males dark



Rana sylvatica (Wood frog)
Family Ranidae

Morphological characteristics:
- body fairly stout with narrow waist
- hind limbs moderately long
- extensive webbing on hind feet
- no disks on toes

Living specimens:
- color tan, brown, gray, or reddish
- distinct dark “mask” around eyes



Rana pipiens (Leopard frog)
Family Ranidae

Morphological characteristics:
- body elongate with slightly-pointed snout, narrow waist
- limbs long and slender
- skin very smooth
- distinct dorsolateral folds extending from the eye to the pelvic region and marked with light stripe
- vocal sacs lateral, located between tympanum and arm

Living specimens:
- dorsal color brownish, grayish, or green
- 2-3 rows of irregular rounded dark spots with light borders
- venter white
- snout usually with a spot, light line on upper lip
- throats of males not dark



Rana palustris (Pickerel frog)
Family Ranidae

Morphological characteristics:
- very similar to R. pipiens
- body elongate with slightly-pointed snout, narrow waist
- limbs long and slender
- skin very smooth
- distinct dorsolateral folds extending from the eye to the pelvic region and marked with light stripe
- vocal sacs lateral, located between tympanum and arm

Living specimens:
- dark dorsal spots are square and dark-bordered
- spots regularly arranged in two rows down back
- square spots present on sides
- overall color tan or brownish, never green
- orange wash on undersides of legs, in groin, and on sides of venter



Rana clamitans (Green frog)
Family Ranidae

Morphological characteristics:
- body shape similar to other ranids
- stouter than R. pipiens and R. palustris
- dorsolateral fold always present, extending from eye to pelvic region
- webbing reaches 2nd joint of 4th toe, does not reach tip of 5th toe
- tympanum of male much larger than eye, tympanum of female same size as eye
- head is not as wide as R. catesbeiana

Living specimens:
- dorsal color green or olive, usually with little mottling or spotting
- throats of males bright yellow



Rana catesbeiana (Bullfrog)
Family Ranidae

Morphological characteristics:
- body large and heavy
- legs very stout
- no dorsolateral fold along sides of body
- distinct fold from eye curving downward along tympanum
- tympanum of male much larger than eye, tympanum of female same size as eye
- extensive webbing on hind feet
- hind toe webs indented on both sides of 4th toe, 4th toe extends beyond web by about one toe segment

Living specimens:

- body color brown or green, variable mottling
- no light spots on lips or light dorsolateral strips
- throats of males bright yellow



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